Behaviour group

Respiration

Breathing-related behaviours that index metabolic demand, oxygen availability, and acute stress.

Breathing index

Opercular movement rate and regularity.

Mouth movement rate

Rate of mouth opening and closing.

Opercular rate

Gill-cover beat frequency.

Yawning

Wide, slow mouth-gape events.

Coughing

Gill-clearing reflex bursts.